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In this wacky spin on the boy-meets-girl tale, Gilly Noble (Chris Klein) enters a sexually charged, passionate love affair with girl of his dreams Jo Wingfield (Heather Graham). After a happy time together, they soon realize that they are brother and sister and quickly break it off. Jo begins a new life and Gilly is left heartbroken -- until he learns that Jo is not actually his sister. As Gilly travels to meet up with his beloved, he finds out that Jo is engaged to be married to another man; he must put a stop to it, even while everyone still believes he is Jo's sibling.